After joining the Federation, the player has to choose one of three career paths. Don't sweat this too much, because any type of officer can fly any type of ship in STO. When I got started, I assumed that making a Science Officer meant I might be stuck with Science Vessels. That's not the case. For our purposes, the main difference among career paths is the type of ground combat abilities you'll receive. While everyone gets various buffs and debuffs, each career has some unique properties.
* Engineering Officer - These are the Scottys and the Chief O'Briens of the world. They're the guys in the yellow shirts shouting about not having the power. (Or was that Ace Ventura?) In STO, Engineers are known for fabrication abilities. They can make turrets that spew grenades, emit energy shields or heal allies.
* Science Officer - These are your Bones McCoys and (sigh) Julian Bashirs. They're the folks in blue shirts who are doctors, not physicists (or escalators, or moon-shuttle conductors.) In STO, Science Officers get most of the healing abilities, as well as some nifty ways to stun, control and otherwise harass enemies.
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